education and training
- Ph.D. in Ecology and Evolutionary Biology, University of California, Irvine , Ecology and Evolutionary Biology, Thesis title: "The dimensions of floral herbivory: Direct and indirect effects of Meligethes rufimanus on the male and female reproductive success of Isomeris arborea" 1990 - 1996
- B.A. in Ecology, Behavior and Evolutionary Biology, University of California, San Diego , Biological Sciences 1986 - 1990