Zhang, Liang, Rothfels, Carl J., Ebihara, Atsushi, Schuettpelz, Eric, Le Pechon, Timothee, Kamau, Peris, He, Hai, Zhou, Xin-Mao, Prado, Jefferson, Field, Ashley, Yatskievych, George, Gao, Xin-Fen, and Zhang, Li-Bing. 2015. "A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae." Cladistics, 31, (4) 406–423. https://doi.org/10.1111/cla.12094.